Except in certain exceptions, most fees paid to Ledgerama for specified services in Wallawallet are paid from the account specified in Settings => USAGE FEES => Payment Account. 

By default, when you create your first account in Wallawallet, this payment account will be automatically defaulted for you.

For different actions you can perform throughout Wallawallet, you will be notified whether or not the service requires fees and from which account they will be paid from. If none has been chosen, you will need to select a payment account before using a service that requires it.

You can also specify the maximum fees you want to pay, either Wallawallet fees or Hedera fees.